diff --git a/csharp/src/Google.Protobuf.Test/WellKnownTypes/WrappersTest.cs b/csharp/src/Google.Protobuf.Test/WellKnownTypes/WrappersTest.cs
index 8a9c3d0c68..4a425f79aa 100644
--- a/csharp/src/Google.Protobuf.Test/WellKnownTypes/WrappersTest.cs
+++ b/csharp/src/Google.Protobuf.Test/WellKnownTypes/WrappersTest.cs
@@ -386,7 +386,7 @@ namespace Google.Protobuf.WellKnownTypes
         }
 
         [Test]
-        public void UnknownFieldInWrapper()
+        public void UnknownFieldInWrapperInt32FastPath()
         {
             var stream = new MemoryStream();
             var output = new CodedOutputStream(stream);
@@ -395,19 +395,96 @@ namespace Google.Protobuf.WellKnownTypes
             var valueTag = WireFormat.MakeTag(Int32Value.ValueFieldNumber, WireFormat.WireType.Varint);
 
             output.WriteTag(wrapperTag);
-            output.WriteLength(4); // unknownTag + value 5 + valueType + value 6, each 1 byte
+            // Wrapper message is just long enough - 6 bytes - to use the wrapper fast-path.
+            output.WriteLength(6); // unknownTag + value 5 + valueType, each 1 byte, + value 65536, 3 bytes
             output.WriteTag(unknownTag);
             output.WriteInt32((int) valueTag); // Sneakily "pretend" it's a tag when it's really a value
             output.WriteTag(valueTag);
+            output.WriteInt32(65536);
+            
+            output.Flush();
+            Assert.AreEqual(8, stream.Length); // tag (1 byte) + length (1 byte) + message (6 bytes)
+            stream.Position = 0;
+
+            var message = TestWellKnownTypes.Parser.ParseFrom(stream);
+            Assert.AreEqual(65536, message.Int32Field);
+        }
+
+        [Test]
+        public void UnknownFieldInWrapperInt32SlowPath()
+        {
+            var stream = new MemoryStream();
+            var output = new CodedOutputStream(stream);
+            var wrapperTag = WireFormat.MakeTag(TestWellKnownTypes.Int32FieldFieldNumber, WireFormat.WireType.LengthDelimited);
+            var unknownTag = WireFormat.MakeTag(15, WireFormat.WireType.Varint);
+            var valueTag = WireFormat.MakeTag(Int32Value.ValueFieldNumber, WireFormat.WireType.Varint);
+
+            output.WriteTag(wrapperTag);
+            // Wrapper message is too short to be used on the wrapper fast-path.
+            output.WriteLength(4); // unknownTag + value 5 + valueType + value 6, each 1 byte
+            output.WriteTag(unknownTag);
+            output.WriteInt32((int)valueTag); // Sneakily "pretend" it's a tag when it's really a value
+            output.WriteTag(valueTag);
             output.WriteInt32(6);
 
             output.Flush();
+            Assert.Less(stream.Length, 8); // tag (1 byte) + length (1 byte) + message
             stream.Position = 0;
 
             var message = TestWellKnownTypes.Parser.ParseFrom(stream);
             Assert.AreEqual(6, message.Int32Field);
         }
 
+        [Test]
+        public void UnknownFieldInWrapperInt64FastPath()
+        {
+            var stream = new MemoryStream();
+            var output = new CodedOutputStream(stream);
+            var wrapperTag = WireFormat.MakeTag(TestWellKnownTypes.Int64FieldFieldNumber, WireFormat.WireType.LengthDelimited);
+            var unknownTag = WireFormat.MakeTag(15, WireFormat.WireType.Varint);
+            var valueTag = WireFormat.MakeTag(Int64Value.ValueFieldNumber, WireFormat.WireType.Varint);
+
+            output.WriteTag(wrapperTag);
+            // Wrapper message is just long enough - 10 bytes - to use the wrapper fast-path.
+            output.WriteLength(11); // unknownTag + value 5 + valueType, each 1 byte, + value 0xfffffffffffff, 8 bytes
+            output.WriteTag(unknownTag);
+            output.WriteInt64((int)valueTag); // Sneakily "pretend" it's a tag when it's really a value
+            output.WriteTag(valueTag);
+            output.WriteInt64(0xfffffffffffffL);
+
+            output.Flush();
+            Assert.AreEqual(13, stream.Length); // tag (1 byte) + length (1 byte) + message (11 bytes)
+            stream.Position = 0;
+
+            var message = TestWellKnownTypes.Parser.ParseFrom(stream);
+            Assert.AreEqual(0xfffffffffffffL, message.Int64Field);
+        }
+
+        [Test]
+        public void UnknownFieldInWrapperInt64SlowPath()
+        {
+            var stream = new MemoryStream();
+            var output = new CodedOutputStream(stream);
+            var wrapperTag = WireFormat.MakeTag(TestWellKnownTypes.Int64FieldFieldNumber, WireFormat.WireType.LengthDelimited);
+            var unknownTag = WireFormat.MakeTag(15, WireFormat.WireType.Varint);
+            var valueTag = WireFormat.MakeTag(Int64Value.ValueFieldNumber, WireFormat.WireType.Varint);
+
+            output.WriteTag(wrapperTag);
+            // Wrapper message is too short to be used on the wrapper fast-path.
+            output.WriteLength(4); // unknownTag + value 5 + valueType + value 6, each 1 byte
+            output.WriteTag(unknownTag);
+            output.WriteInt64((int)valueTag); // Sneakily "pretend" it's a tag when it's really a value
+            output.WriteTag(valueTag);
+            output.WriteInt64(6);
+
+            output.Flush();
+            Assert.Less(stream.Length, 12); // tag (1 byte) + length (1 byte) + message
+            stream.Position = 0;
+
+            var message = TestWellKnownTypes.Parser.ParseFrom(stream);
+            Assert.AreEqual(6L, message.Int64Field);
+        }
+
         [Test]
         public void ClearWithReflection()
         {
diff --git a/csharp/src/Google.Protobuf/CodedInputStream.cs b/csharp/src/Google.Protobuf/CodedInputStream.cs
index 4d13f88ab9..44934f341b 100644
--- a/csharp/src/Google.Protobuf/CodedInputStream.cs
+++ b/csharp/src/Google.Protobuf/CodedInputStream.cs
@@ -481,7 +481,33 @@ namespace Google.Protobuf
         /// </summary>
         public double ReadDouble()
         {
-            return BitConverter.Int64BitsToDouble((long) ReadRawLittleEndian64());
+            if (bufferPos + 8 <= bufferSize)
+            {
+                if (BitConverter.IsLittleEndian)
+                {
+                    var result = BitConverter.ToDouble(buffer, bufferPos);
+                    bufferPos += 8;
+                    return result;
+                }
+                else
+                {
+                    var bytes = new byte[8];
+                    bytes[0] = buffer[bufferPos + 7];
+                    bytes[1] = buffer[bufferPos + 6];
+                    bytes[2] = buffer[bufferPos + 5];
+                    bytes[3] = buffer[bufferPos + 4];
+                    bytes[4] = buffer[bufferPos + 3];
+                    bytes[5] = buffer[bufferPos + 2];
+                    bytes[6] = buffer[bufferPos + 1];
+                    bytes[7] = buffer[bufferPos];
+                    bufferPos += 8;
+                    return BitConverter.ToDouble(bytes, 0);
+                }
+            }
+            else
+            {
+                return BitConverter.Int64BitsToDouble((long)ReadRawLittleEndian64());
+            }
         }
 
         /// <summary>
@@ -711,7 +737,260 @@ namespace Google.Protobuf
             return false;
         }
 
-        #endregion
+        internal static float? ReadFloatWrapperLittleEndian(CodedInputStream input)
+        {
+            // length:1 + tag:1 + value:4 = 6 bytes
+            if (input.bufferPos + 6 <= input.bufferSize)
+            {
+                // The entire wrapper message is already contained in `buffer`.
+                int length = input.buffer[input.bufferPos];
+                if (length == 0)
+                {
+                    input.bufferPos++;
+                    return 0F;
+                }
+                // tag:1 + value:4 = length of 5 bytes
+                // field=1, type=32-bit = tag of 13
+                if (length != 5 || input.buffer[input.bufferPos + 1] != 13)
+                {
+                    return ReadFloatWrapperSlow(input);
+                }
+                var result = BitConverter.ToSingle(input.buffer, input.bufferPos + 2);
+                input.bufferPos += 6;
+                return result;
+            }
+            else
+            {
+                return ReadFloatWrapperSlow(input);
+            }
+        }
+
+        internal static float? ReadFloatWrapperSlow(CodedInputStream input)
+        {
+            int length = input.ReadLength();
+            if (length == 0)
+            {
+                return 0F;
+            }
+            int finalBufferPos = input.totalBytesRetired + input.bufferPos + length;
+            float result = 0F;
+            do
+            {
+                // field=1, type=32-bit = tag of 13
+                if (input.ReadTag() == 13)
+                {
+                    result = input.ReadFloat();
+                }
+                else
+                {
+                    input.SkipLastField();
+                }
+            }
+            while (input.totalBytesRetired + input.bufferPos < finalBufferPos);
+            return result;
+        }
+
+        internal static double? ReadDoubleWrapperLittleEndian(CodedInputStream input)
+        {
+            // length:1 + tag:1 + value:8 = 10 bytes
+            if (input.bufferPos + 10 <= input.bufferSize)
+            {
+                // The entire wrapper message is already contained in `buffer`.
+                int length = input.buffer[input.bufferPos];
+                if (length == 0)
+                {
+                    input.bufferPos++;
+                    return 0D;
+                }
+                // tag:1 + value:8 = length of 9 bytes
+                // field=1, type=64-bit = tag of 9
+                if (length != 9 || input.buffer[input.bufferPos + 1] != 9)
+                {
+                    return ReadDoubleWrapperSlow(input);
+                }
+                var result = BitConverter.ToDouble(input.buffer, input.bufferPos + 2);
+                input.bufferPos += 10;
+                return result;
+            }
+            else
+            {
+                return ReadDoubleWrapperSlow(input);
+            }
+        }
+
+        internal static double? ReadDoubleWrapperSlow(CodedInputStream input)
+        {
+            int length = input.ReadLength();
+            if (length == 0)
+            {
+                return 0D;
+            }
+            int finalBufferPos = input.totalBytesRetired + input.bufferPos + length;
+            double result = 0D;
+            do
+            {
+                // field=1, type=64-bit = tag of 9
+                if (input.ReadTag() == 9)
+                {
+                    result = input.ReadDouble();
+                }
+                else
+                {
+                    input.SkipLastField();
+                }
+            }
+            while (input.totalBytesRetired + input.bufferPos < finalBufferPos);
+            return result;
+        }
+
+        internal static bool? ReadBoolWrapper(CodedInputStream input)
+        {
+            return ReadUInt32Wrapper(input) != 0;
+        }
+
+        internal static uint? ReadUInt32Wrapper(CodedInputStream input)
+        {
+            // length:1 + tag:1 + value:5(varint32-max) = 7 bytes
+            if (input.bufferPos + 7 <= input.bufferSize)
+            {
+                // The entire wrapper message is already contained in `buffer`.
+                int pos0 = input.bufferPos;
+                int length = input.buffer[input.bufferPos++];
+                if (length == 0)
+                {
+                    return 0;
+                }
+                // Length will always fit in a single byte.
+                if (length >= 128)
+                {
+                    input.bufferPos = pos0;
+                    return ReadUInt32WrapperSlow(input);
+                }
+                int finalBufferPos = input.bufferPos + length;
+                // field=1, type=varint = tag of 8
+                if (input.buffer[input.bufferPos++] != 8)
+                {
+                    input.bufferPos = pos0;
+                    return ReadUInt32WrapperSlow(input);
+                }
+                var result = input.ReadUInt32();
+                // Verify this message only contained a single field.
+                if (input.bufferPos != finalBufferPos)
+                {
+                    input.bufferPos = pos0;
+                    return ReadUInt32WrapperSlow(input);
+                }
+                return result;
+            }
+            else
+            {
+                return ReadUInt32WrapperSlow(input);
+            }
+        }
+
+        private static uint? ReadUInt32WrapperSlow(CodedInputStream input)
+        {
+            int length = input.ReadLength();
+            if (length == 0)
+            {
+                return 0;
+            }
+            int finalBufferPos = input.totalBytesRetired + input.bufferPos + length;
+            uint result = 0;
+            do
+            {
+                // field=1, type=varint = tag of 8
+                if (input.ReadTag() == 8)
+                {
+                    result = input.ReadUInt32();
+                }
+                else
+                {
+                    input.SkipLastField();
+                }
+            }
+            while (input.totalBytesRetired + input.bufferPos < finalBufferPos);
+            return result;
+        }
+
+        internal static int? ReadInt32Wrapper(CodedInputStream input)
+        {
+            return (int?)ReadUInt32Wrapper(input);
+        }
+
+        internal static ulong? ReadUInt64Wrapper(CodedInputStream input)
+        {
+            // field=1, type=varint = tag of 8
+            const int expectedTag = 8;
+            // length:1 + tag:1 + value:10(varint64-max) = 12 bytes
+            if (input.bufferPos + 12 <= input.bufferSize)
+            {
+                // The entire wrapper message is already contained in `buffer`.
+                int pos0 = input.bufferPos;
+                int length = input.buffer[input.bufferPos++];
+                if (length == 0)
+                {
+                    return 0L;
+                }
+                // Length will always fit in a single byte.
+                if (length >= 128)
+                {
+                    input.bufferPos = pos0;
+                    return ReadUInt64WrapperSlow(input);
+                }
+                int finalBufferPos = input.bufferPos + length;
+                if (input.buffer[input.bufferPos++] != expectedTag)
+                {
+                    input.bufferPos = pos0;
+                    return ReadUInt64WrapperSlow(input);
+                }
+                var result = input.ReadUInt64();
+                // Verify this message only contained a single field.
+                if (input.bufferPos != finalBufferPos)
+                {
+                    input.bufferPos = pos0;
+                    return ReadUInt64WrapperSlow(input);
+                }
+                return result;
+            }
+            else
+            {
+                return ReadUInt64WrapperSlow(input);
+            }
+        }
+
+        internal static ulong? ReadUInt64WrapperSlow(CodedInputStream input)
+        {
+            // field=1, type=varint = tag of 8
+            const int expectedTag = 8;
+            int length = input.ReadLength();
+            if (length == 0)
+            {
+                return 0L;
+            }
+            int finalBufferPos = input.totalBytesRetired + input.bufferPos + length;
+            ulong result = 0L;
+            do
+            {
+                if (input.ReadTag() == expectedTag)
+                {
+                    result = input.ReadUInt64();
+                }
+                else
+                {
+                    input.SkipLastField();
+                }
+            }
+            while (input.totalBytesRetired + input.bufferPos < finalBufferPos);
+            return result;
+        }
+
+        internal static long? ReadInt64Wrapper(CodedInputStream input)
+        {
+            return (long?)ReadUInt64Wrapper(input);
+        }
+
+#endregion
 
         #region Underlying reading primitives
 
@@ -876,17 +1155,42 @@ namespace Google.Protobuf
         /// </summary>
         internal ulong ReadRawVarint64()
         {
-            int shift = 0;
-            ulong result = 0;
-            while (shift < 64)
+            if (bufferPos + 10 <= bufferSize)
             {
-                byte b = ReadRawByte();
-                result |= (ulong) (b & 0x7F) << shift;
-                if ((b & 0x80) == 0)
+                ulong result = buffer[bufferPos++];
+                if (result < 128)
                 {
                     return result;
                 }
-                shift += 7;
+                result &= 0x7f;
+                int shift = 7;
+                do
+                {
+                    byte b = buffer[bufferPos++];
+                    result |= (ulong)(b & 0x7F) << shift;
+                    if (b < 0x80)
+                    {
+                        return result;
+                    }
+                    shift += 7;
+                }
+                while (shift < 64);
+            }
+            else
+            {
+                int shift = 0;
+                ulong result = 0;
+                do
+                {
+                    byte b = ReadRawByte();
+                    result |= (ulong)(b & 0x7F) << shift;
+                    if (b < 0x80)
+                    {
+                        return result;
+                    }
+                    shift += 7;
+                }
+                while (shift < 64);
             }
             throw InvalidProtocolBufferException.MalformedVarint();
         }
@@ -896,11 +1200,32 @@ namespace Google.Protobuf
         /// </summary>
         internal uint ReadRawLittleEndian32()
         {
-            uint b1 = ReadRawByte();
-            uint b2 = ReadRawByte();
-            uint b3 = ReadRawByte();
-            uint b4 = ReadRawByte();
-            return b1 | (b2 << 8) | (b3 << 16) | (b4 << 24);
+            if (bufferPos + 4 <= bufferSize)
+            {
+                if (BitConverter.IsLittleEndian)
+                {
+                    var result = BitConverter.ToUInt32(buffer, bufferPos);
+                    bufferPos += 4;
+                    return result;
+                }
+                else
+                {
+                    uint b1 = buffer[bufferPos];
+                    uint b2 = buffer[bufferPos + 1];
+                    uint b3 = buffer[bufferPos + 2];
+                    uint b4 = buffer[bufferPos + 3];
+                    bufferPos += 4;
+                    return b1 | (b2 << 8) | (b3 << 16) | (b4 << 24);
+                }
+            }
+            else
+            {
+                uint b1 = ReadRawByte();
+                uint b2 = ReadRawByte();
+                uint b3 = ReadRawByte();
+                uint b4 = ReadRawByte();
+                return b1 | (b2 << 8) | (b3 << 16) | (b4 << 24);
+            }
         }
 
         /// <summary>
@@ -908,16 +1233,42 @@ namespace Google.Protobuf
         /// </summary>
         internal ulong ReadRawLittleEndian64()
         {
-            ulong b1 = ReadRawByte();
-            ulong b2 = ReadRawByte();
-            ulong b3 = ReadRawByte();
-            ulong b4 = ReadRawByte();
-            ulong b5 = ReadRawByte();
-            ulong b6 = ReadRawByte();
-            ulong b7 = ReadRawByte();
-            ulong b8 = ReadRawByte();
-            return b1 | (b2 << 8) | (b3 << 16) | (b4 << 24)
-                   | (b5 << 32) | (b6 << 40) | (b7 << 48) | (b8 << 56);
+            if (bufferPos + 8 <= bufferSize)
+            {
+                if (BitConverter.IsLittleEndian)
+                {
+                    var result = BitConverter.ToUInt64(buffer, bufferPos);
+                    bufferPos += 8;
+                    return result;
+                }
+                else
+                {
+                    ulong b1 = buffer[bufferPos];
+                    ulong b2 = buffer[bufferPos + 1];
+                    ulong b3 = buffer[bufferPos + 2];
+                    ulong b4 = buffer[bufferPos + 3];
+                    ulong b5 = buffer[bufferPos + 4];
+                    ulong b6 = buffer[bufferPos + 5];
+                    ulong b7 = buffer[bufferPos + 6];
+                    ulong b8 = buffer[bufferPos + 7];
+                    bufferPos += 8;
+                    return b1 | (b2 << 8) | (b3 << 16) | (b4 << 24)
+                           | (b5 << 32) | (b6 << 40) | (b7 << 48) | (b8 << 56);
+                }
+            }
+            else
+            {
+                ulong b1 = ReadRawByte();
+                ulong b2 = ReadRawByte();
+                ulong b3 = ReadRawByte();
+                ulong b4 = ReadRawByte();
+                ulong b5 = ReadRawByte();
+                ulong b6 = ReadRawByte();
+                ulong b7 = ReadRawByte();
+                ulong b8 = ReadRawByte();
+                return b1 | (b2 << 8) | (b3 << 16) | (b4 << 24)
+                       | (b5 << 32) | (b6 << 40) | (b7 << 48) | (b8 << 56);
+            }
         }
 
         /// <summary>
@@ -1301,6 +1652,6 @@ namespace Google.Protobuf
                 }
             }
         }
-        #endregion
+#endregion
     }
 }
\ No newline at end of file
diff --git a/csharp/src/Google.Protobuf/FieldCodec.cs b/csharp/src/Google.Protobuf/FieldCodec.cs
index ebd00b7022..1971261649 100644
--- a/csharp/src/Google.Protobuf/FieldCodec.cs
+++ b/csharp/src/Google.Protobuf/FieldCodec.cs
@@ -507,7 +507,7 @@ namespace Google.Protobuf
         {
             var nestedCodec = WrapperCodecs.GetCodec<T>();
             return new FieldCodec<T?>(
-                input => WrapperCodecs.Read<T>(input, nestedCodec),
+                WrapperCodecs.GetReader<T>(),
                 (output, value) => WrapperCodecs.Write<T>(output, value.Value, nestedCodec),
                 (CodedInputStream i, ref T? v) => v = WrapperCodecs.Read<T>(i, nestedCodec),
                 (ref T? v, T? v2) => { if (v2.HasValue) { v = v2; } return v.HasValue; },
@@ -539,6 +539,25 @@ namespace Google.Protobuf
                 { typeof(ByteString), ForBytes(WireFormat.MakeTag(WrappersReflection.WrapperValueFieldNumber, WireFormat.WireType.LengthDelimited)) }
             };
 
+            private static readonly Dictionary<System.Type, object> Readers = new Dictionary<System.Type, object>
+            {
+                // TODO: Provide more optimized readers.
+                { typeof(bool), (Func<CodedInputStream, bool?>)CodedInputStream.ReadBoolWrapper },
+                { typeof(int), (Func<CodedInputStream, int?>)CodedInputStream.ReadInt32Wrapper },
+                { typeof(long), (Func<CodedInputStream, long?>)CodedInputStream.ReadInt64Wrapper },
+                { typeof(uint), (Func<CodedInputStream, uint?>)CodedInputStream.ReadUInt32Wrapper },
+                { typeof(ulong), (Func<CodedInputStream, ulong?>)CodedInputStream.ReadUInt64Wrapper },
+                { typeof(float), BitConverter.IsLittleEndian ?
+                    (Func<CodedInputStream, float?>)CodedInputStream.ReadFloatWrapperLittleEndian :
+                    (Func<CodedInputStream, float?>)CodedInputStream.ReadFloatWrapperSlow },
+                { typeof(double), BitConverter.IsLittleEndian ?
+                    (Func<CodedInputStream, double?>)CodedInputStream.ReadDoubleWrapperLittleEndian :
+                    (Func<CodedInputStream, double?>)CodedInputStream.ReadDoubleWrapperSlow },
+                // `string` and `ByteString` less performance-sensitive. Do not implement for now.
+                { typeof(string), null },
+                { typeof(ByteString), null },
+            };
+
             /// <summary>
             /// Returns a field codec which effectively wraps a value of type T in a message.
             ///
@@ -553,6 +572,23 @@ namespace Google.Protobuf
                 return (FieldCodec<T>) value;
             }
 
+            internal static Func<CodedInputStream, T?> GetReader<T>() where T : struct
+            {
+                object value;
+                if (!Readers.TryGetValue(typeof(T), out value))
+                {
+                    throw new InvalidOperationException("Invalid type argument requested for wrapper reader: " + typeof(T));
+                }
+                if (value == null)
+                {
+                    // Return default unoptimized reader for the wrapper type.
+                    var nestedCoded = GetCodec<T>();
+                    return input => Read<T>(input, nestedCoded);
+                }
+                // Return optimized read for the wrapper type.
+                return (Func<CodedInputStream, T?>)value;
+            }
+
             internal static T Read<T>(CodedInputStream input, FieldCodec<T> codec)
             {
                 int length = input.ReadLength();